Ownership Submission
FORM 3
 UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

INITIAL STATEMENT OF BENEFICIAL OWNERSHIP OF SECURITIES

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940
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(Print or Type Responses)
1. Name and Address of Reporting Person *
  Stevenson Mark
 2. Date of Event Requiring Statement (Month/Day/Year)
01/18/2007		 3. Issuer Name and Ticker or Trading Symbol
APPLERA CORP [ABI/CRA]
(Last)
(First)
(Middle)
APPLERA CORP - APPLIED BIOSYSTEMS GROUP, 850 LINCOLN CENTRE DRIVE		 4. Relationship of Reporting Person(s) to Issuer
(Check all applicable)
_____ Director _____ 10% Owner
__X__ Officer (give title below) _____ Other (specify below)
Vice President
 5. If Amendment, Date Original Filed(Month/Day/Year)
(Street)

FOSTER CITY, CA 94404		 6. Individual or Joint/Group Filing(Check Applicable Line)
_X_ Form filed by One Reporting Person
___ Form filed by More than One Reporting Person
(City)
(State)
(Zip)
Table I - Non-Derivative Securities Beneficially Owned
1.Title of Security
(Instr. 4)		 2. Amount of Securities Beneficially Owned
(Instr. 4)		 3. Ownership Form: Direct (D) or Indirect (I)
(Instr. 5)		 4. Nature of Indirect Beneficial Ownership
(Instr. 5)
Applied Biosystems Group Common Stock 0
D
 
Celera Group Common Stock 1,542
D
 

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly. SEC 1473 (7-02)
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Table II - Derivative Securities Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security
(Instr. 4)		 2. Date Exercisable and Expiration Date
(Month/Day/Year)		 3. Title and Amount of Securities Underlying Derivative Security
(Instr. 4)		 4. Conversion or Exercise Price of Derivative Security 5. Ownership Form of Derivative Security: Direct (D) or Indirect (I)
(Instr. 5)		 6. Nature of Indirect Beneficial Ownership
(Instr. 5)
Date Exercisable Expiration Date Title Amount or Number of Shares
Applied Biosystems Group Employee Stock Options-Right to Buy 04/13/2004 04/13/2010 Applied Biosystems Group Common Stock 10,000 $ 81 D  
Applied Biosystems Group Employee Stock Options-Right to Buy   (1) 03/24/2013 Applied Biosystems Group Common Stock 2,875 $ 15.54 D  
Applied Biosystems Group Employee Stock Options-Right to Buy   (2) 03/24/2013 Applied Biosystems Group Common Stock 5,000 $ 15.54 D  
Applied Biosystems Group Employee Stock Options-Right to Buy   (3) 06/17/2014 Applied Biosystems Group Common Stock 7,500 $ 20.185 D  
Applied Biosystems Group Employee Stock Options-Right to Buy 06/02/2005(4) 06/02/2015 Applied Biosystems Group Common Stock 45,000 $ 21.475 D  

Reporting Owners

Reporting Owner Name / Address Relationships
Director 10% Owner Officer Other
Stevenson Mark
APPLERA CORP - APPLIED BIOSYSTEMS GROUP
850 LINCOLN CENTRE DRIVE
FOSTER CITY, CA 94404		       Vice President  

Signatures

/s/ Thomas P. Livingston, Attorney-In-Fact for Mark Stevenson 01/29/2007
**Signature of Reporting Person Date

Explanation of Responses:

* If the form is filed by more than one reporting person, see Instruction 5(b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
(1) These options originally became exercisable in four equal annual installments, commencing March 24, 2004. Effective January 20, 2005, the vesting of all of these options was accelerated. However, shares received upon the exercise of accelerated options are subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition) that will remain in effect until the earlier of the original vesting date of those options or termination of employment.
(2) As originally granted, these options would have become exercisable on March 24, 2008. Effective June 2, 2005, the vesting of all of these options was accelerated. However, shares received upon the exercise of accelerated options are subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition) that will remain in effect until the earlier of the vesting date of those options, i.e. March 24, 2008, or termination of employment.
(3) These options originally became exercisable in four equal annual installments, commencing June 17, 2005. Effective January 20, 2005, the vesting of all of these options was accelerated. However, shares received upon the exercise of accelerated options are subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition) that will remain in effect until the earlier of the original vesting date of those options or termination of employment.
(4) These options are fully vested and exercisable as of June 2, 2005, the grant date. However, shares acquired upon the exercise of these options are subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition). The transfer restriction will lapse on 25% of the shares covered by the grant on each of the first four anniversaries of the grant date. Also, the transfer restriction will lapse in full upon termination of employment.

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